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NEUBEH 501 Introduction to Neurobiology (3) Carlson
Survey of molecular, cellular, and developmental neuroscience, including gene regulation, the cytoskeleton, protein sorting in the secretory pathway, growth factors, and neurotransmitter receptors. Includes lecture discussion of original literature. Offered: A.

NEUBEH 502 Introduction to Neurobiology (5) Sherk
Systems level survey of vertebrate system, focusing on sensory system, on motor system, and on neuroanatomy. Lectures cover topics in sensory and motor systems. Laboratory includes brain dissection and study of intact, prosected, and sectioned brain and spinal cord. Emphasis is on human nervous system. Offered: W.

NEUBEH 503 Cognitive and Integrative Neurobiology (4) Phillips
Survey of all aspects of neuroscience, including a discussion of higher neural processes like motivation, decision making, attention, learning, and memory. Lecture and discussion of original literature. Offered: Sp.

NEUBEH 504 Biophysics of Nerve, Muscle, and Synapse (3) Sullivan
Introduces biophysical properties of nerve and muscle cells. Topics include intrinsic electrical properties of neurons, ion channels, receptor signaling, calcium signaling, contraction of muscles, and synaptic function. Offered: jointly with P BIO 504; A.

NEUBEH 510 Seminar in Neurobiology and Behavior (0.5, max. 30)
Biweekly seminar on current topics. Required for students in the graduate program in neurobiology and behavior and for students supported on Graduate Neuroscience Program Training Grant. Credit/no-credit only. Offered: AWSp.

NEUBEH 511 Seminar in Advanced Neurobiology (1-3, max. 30)
Weekly faculty lectures, student presentations, and discussions of past and current scientific literature in neurobiology and behavior. Offered: AWSpS.

NEUBEH 512 Readings in Advanced Neurobiology and Behavior (1-3, max. 30)
Guided study of the primary literature of neurobiology and behavior. Emphasizes critical analysis, accuracy of expression, bibliographical technique, and other factors of good scholarship. Prerequisite: permission of instructor. Credit/no-credit only. Offered: AWSpS.

NEUBEH 515 Teaching Practicum in Neurobiology and Behavior (3-6, max. 30)
Supervised training in the teaching of neuroscience and related scientific topics. Prerequisite: graduate standing in the neurobiology and behavior graduate program and permission of the instructor. Credit/no-credit only. Offered: AWSpS.

NEUBEH 526 Introduction to Laboratory Research in Neurobiology (4, max. 30)
Students become familiar with, and assist in, the performance of research on ongoing projects in designated laboratories. Emphasis on employed methodology and techniques. Prerequisite: first-year graduate students in neurobiology. Credit/no-credit only. Offered: AWSpS.

NEUBEH 527 Current Topics in Neurobiology and Behavior (1, max. 30) Sullivan
Presentation and discussion of current research provides exposure to diverse areas of neurobiology and behavior research. Prerequisite: graduate student in neurobiology and behavior program or permission of instructor. Credit/no-credit only. Offered: AWSp.

NEUBEH 528 Computational Neuroscience (3)
Introduction to computational methods for understanding nervous systems and the principles governing their operation. Topics include representation of information by spiking neurons, information processing in neural circuits, and algorithms for adaptation and learning. Prerequisite: elementary calculus, linear algebra, and statistics, or permission of instructor. Offered: jointly with CSE 528.

NEUBEH 532 Discussion in Cell Signaling and Molecular Physiology (3) Fairhall, Rieke
Discusses fundamental issues in cell excitability and molecular and cellular physiology. Focuses on problem solving and reading from original literature. Emphasizes student participation. Prerequisite: first-year graduate students in neurobiology or physiology and biophysics. Offered: jointly with P BIO 532; W.

NEUBEH 541 Neuroendocrinology (3) Steiner
Emphasizes the cellular and molecular aspects of several topics in neuroendocrinology, including neuropeptide genes, reproduction, steroid hormone regulation of gene expression, mechanisms of hormone action, endocrine rhythms, and neural oscillations. Prerequisite: either BIOL 201, BIOL 202, and BIOL 203, or BIOL 180, BIOL 200, and BIOL 220; BIOC 440, BIOC 441, BIOC 442 or permission of instructor. Offered: jointly with P BIO 509; W, odd years.

NEUBEH 545 Quantitative Methods in Neuroscience (3) Rieke, Shadlen
Discusses quantitative methods applicable to the study of the nervous system. Revolves around computer exercises/discussion of journal papers. May include linear systems theory, Fourier analysis, ordinary differential equations, stochastic processes, signal detection, and information theory. Prerequisite: NEUBEH 501, NEUBEH 502, NEUBEH 503, or permission of instructor. Offered: jointly with P BIO 545; W, odd years.

NEUBEH 548 Molecular Mechanisms of Synaptic Plasticity (2) Barria
Discusses recent primary literature on the molecular mechanisms underlying structural and functional changes of dendritic spines and synapses in the mammalian brain as result of synaptic activity and experience. Offered: jointly with P BIO 548; Sp, odd years.

NEUBEH 549 Molecular Basis of Neurodegenerative Disease (2) Pallanck
Introduces a broad range of neurodegenerative diseases, focusing upon the approaches that have led to recent discoveries and emphasizing the elucidation of mechanisms and pathways of disease pathogenesis. Offered: jointly with GENOME 549/PHCOL 549; W.

NEUBEH 550 Biophysics of Calcium Signaling (1) Hille, Santana
Introduction to cellular calcium signaling including theoretical and technical issues of calcium signal detection and biological conclusions. Prerequisite: CONJ 531. Offered: jointly with P BIO 550; Sp, odd years.

NEUBEH 552 Synaptic Integration (1) Binder, Powers
Discussion of recent papers on how neurons in the central nervous system integrate concurrent synaptic inputs. Includes: effects of driving force on synaptic currents, effects of conductances on dendritic properties, transfer of currents from dendrites to soma, and transformation of currents into spike train outputs. Offered: jointly with P BIO 552.

NEUBEH 553 Learning and Memory: Synapses and Systems (2) Jagadeesh, Sullivan
Five-week mini-course evaluates the current state of knowledge on the mechanisms that allow people to learn and remember. After introductory overviews of the cellular and molecular mechanisms underlying long-term synaptic plasticity and the multiple systems existing for learning and memory, students choose specific topics for discussion. Offered: jointly with P BIO 553.

NEUBEH 554 Motor Learning: Cellular and Network Mechanisms (1) Fetz, Perlmutter
Five-week mini-course reviews the current state of research on cellular and network mechanisms of motor learning. After an introductory overview of behavioral and physiological examples of motor learning in various species and systems, students choose specific topics for discussion, using the primary literature as a source. Offered: jointly with P BIO 554.
Instructor Course Description: Eberhard Fetz

NEUBEH 555 Sensory Receptors (1) Detwiler, Rieke
Five-lecture mini-course examines how different kinds of sensory receptors detect and respond to different modalities of sensory stimuli. Discussion focuses on the cellular and molecular mechanisms of the underlying transduction processes and the experimental evidence that they are based on. Offered: jointly with P BIO 555.

NEUBEH 556 Axon Pathfinding Mechanisms (1) Bothwell
Examines mechanisms governing axon growth cone behavior during embryonic development and during regeneration in the injured adult. Discusses approaches employing both invertebrate and vertebrate model systems. Offered: jointly with P BIO 556.

NEUBEH 557 Ion Channel Gating (1) Zagotta
Compares and contrasts mechanisms of gating in ligand-gated and voltage-gated ion channels. Covers basics of ligand gating and voltage gating, kinetic schemes, inactivation and desensitization, gating currents and partial agonists, and ion channel structure. Offered: jointly with P BIO 557.

NEUBEH 559 Neurobiology of Disease (3) Garden, Neumaier, Weinstein, Weiss
Introduces medically important neurological and psychiatric diseases and experimental approaches to understanding the basis for diseases and their treatments. Covers stroke, epilespsy, autoimmune diseases of the CNS, neurodegenerative diseases, autism, psychosis, anxiety disorders, and mood disorders. Offered: jointly with NEURL 559/P BIO 559.

NEUBEH 561 From Biophysics to Neural Computation (2)
Introduces the mathematics and methods of neuronal modeling. Develops, compares, and relates dynamical systems approaches and empirical characterizations of neuronal function. Includes lectures, student-led journal paper discussions, biweekly computational workshops on neuronal modeling packages, and a computations project. Recommended: differential equations and linear algebra. Offered: jointly with P BIO 561; Sp.

NEUBEH 600 Independent Study or Research (*-)
Credit/no-credit only. Offered: AWSpS.
Instructor Course Description: O Bermingham-Mcdonogh

NEUBEH 700 Master's Thesis (*-)
Offered: AWSpS.

NEUBEH 800 Doctoral Dissertation (*-)
Offered: AWSpS.