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COLLEGE OF ARTS & SCIENCES
BIOLOGY

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BIOL 100 Introductory Biology (5) NW
Develops an awareness of science by studying basic biological principles and their application to problems of humans and society in the contexts of special topics or themes, which vary quarter to quarter. For non-science majors only.
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BIOL 106 Introductory Biology Seminar (1/3, max. 6) NW
Focuses on current topics in biology. Topics vary from quarter to quarter. Designed to enhance learning skills of students who intend to take BIOL 180/BIOL 200/BIOL 220 and major in one of the biological sciences. Recommended: high school chemistry and biology, or one quarter of college chemistry.
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BIOL 108 Evolution and Human Behavior (3) NW
Introduction to evolution by natural selection, examining the light it can throw on human biology and behavior in such areas as the nature of sex differences, sexual conflict, and conflict between parents and children. Does not fulfill major requirements. Offered: jointly with BIO A 100.
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BIOL 110 Freshman Discovery Seminar in Biology (5) NW
Introduces incoming freshman to research basics and scholarly inquiry skills used in the study of biology.
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BIOL 114 Astrobiology: Life in the Universe (5) NW R. BUICK, D. CATLING, V. MEADOWS, W. SULLIVAN
Introduction to the new science of astrobiology, study of the origin and evolution of life on Earth, and the search for microbial and intelligent life elsewhere in the Universe. Designed for non-science, liberal arts majors. Offered: jointly with ASTBIO 115/ASTR 115/ESS 115/OCEAN 115.
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BIOL 118 Survey of Physiology (5) NW
Human physiology, for nonmajors and health sciences students. Offered: AWSpS.
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BIOL 119 Elementary Physiology Laboratory (1) NW
Prerequisite: BIOL 118 which may be taken concurrently. Offered: AWSpS.
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BIOL 120 Current Controversies in Biology (2-5, max. 6) NW
Explores a current controversial topic in biology, stressing information needed by the general public to make informed personal, political, and ethical decisions relating to this topic.
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BIOL 180 Introductory Biology (5) NW
Mendelian genetics, evolution, biodiversity of life forms, ecology, and conservation biology. Open to all students interested in biology whether intending to major in the biological sciences, enroll in preprofessional programs, or fulfill a Natural World requirement. First course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Offered: AWSpS.
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BIOL 200 Introductory Biology (5) NW
For students intending to take advanced courses in the biological sciences or enroll in preprofessional programs. Metabolism and energetics, structure and function of biomolecules, cell structure and function, animal development. Second course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Prerequisite: minimum grade of 1.7 in either BIOL 180, B BIO 180, or TESC 120; either CHEM 152, CHEM 155, CHEM 220, CHEM 223, or CHEM 237. Offered: AWSpS.
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BIOL 220 Introductory Biology (5) NW
For students intending to take advanced courses in the biological sciences or enroll in preprofessional programs. Animal physiology, plant development and physiology. Final course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Prerequisite: minimum grade of 2.0 in either BIOL 200, B BIO 220, or TESC 130. Offered: AWSpS.
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BIOL 250 Marine Biology (3/5) I&S/NW
Lecture-laboratory course in marine biology focusing on physical, biological, and social aspects of the marine environment. Topics include oceanography, ecology, physiology, behavior, conservation, fisheries, exploration, and activism. Weekend field trip. Honors section research project. Offered: jointly with FISH 250/OCEAN 250; AS.
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BIOL 280 The History of Life (4) NW P. WARD
Follows the history of life from its first formation including the origin of life and life's diversification from single cells through multi-celluarity. Examines fossils and DNA evidence from understanding the sequence of events and evolutionary history of life.
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BIOL 293 Study Abroad - Biology (1-10, max. 10) NW
For participants in UW Study Abroad program. Specific content varies and must be individually evaluated. Credit does not apply to major requirements without approval.
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BIOL 300 Introduction to Neuroscience (4) NW
Provides a broad introduction to the study of brain function in humans and other animals. Emphasizes how circuits within the brain process sensory information and generate complex movements. No credit if NBIO 301, NBIO 302, or BIOL 461 already taken. Prerequisite: either MATH 111, MATH 112, MATH 120, MATH 124, MATH 144, Q SCI 190, or Q SCI 291. Offered: W.
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BIOL 302 Laboratory Techniques in Cell and Molecular Biology (4) NW T. IMAIZUMI, L. MARTIN-MORRIS
Explores the use of various cell and molecular laboratory techniques, such as PCR, cloning, gel electrophoresis, and bacterial transformation through hands-on experiments. Students produce a portfolio of techniques they have learned. Writing credit. Prerequisite: either BIOL 355 or a minimum grade of 2.5 in either BIOL 200, B BIO 200, or TESC 130.
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BIOL 305 Science Communication: Video Storytelling in Biology (3) VLPA/NW P. BOERSMA
Students make a short film on a biological story, concept, or theory. Includes developing a storyline, getting the shots to make compelling viewing, editing, and producing a short video. Prerequisite: either BIOL 180, B BIO 180, or TESC 120. Offered: Sp.
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BIOL 311 Biology of Fishes (3/5) NW
Covers morphological, physiological, behavioral, and ecological diversity of fishes of the world; designed to provide a basic foundation for advanced courses in all areas of aquatic sciences. 3-credit option does not include laboratory. Recommended: 10 credits biological science. Offered: jointly with FISH 311; W.
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BIOL 315 Biological Impacts of Climate Change (3) NW
Covers the biological impacts of climate change, including changes in species distributions and interactions, altered phenology, and ecosystem dynamics. Discusses implications of these biological impacts for society (e.g., food security, public health, and resource management). Prerequisite: either BIOL 180, B BIO 180, or TESC 120.
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BIOL 317 Plant Classification and Identification (5) NW R. OLMSTEAD
Classification and diversity of seed plants; concepts and principles of classification, lab and field study of common plant families in Washington, and skill development for identification of species. One weekend field trip. Offered: SpS.
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BIOL 325 The Tree of Life (4) NW S. FREEMAN
Provides an overview of biodiversity by focusing on key characteristics of major lineage throughout the tree of life. Explores major diversification events by analyzing changes in reproduction, energetics, cell structure, sensory systems, and adaptations to abiotic stress, and species interactions. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
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BIOL 330 Natural History of Marine Invertebrates (5) NW
Field and laboratory course emphasizing the habits, habitats, adaptations, and interrelationships of marine animals. Weekend field trips required. Offered: S.
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BIOL 331 Landscape Plant Recognition (3) NW S. REICHARD
Field recognition of important groups of woody landscape plants, emphasizing diversity at the genus and family levels. Cultivated plant nomenclature. Plant descriptive characters evident in the field with eye and hand lens. Hardiness and landscape applications. Recommended: BIOL 317. Offered: jointly with ESRM 331; Sp.
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BIOL 340 Genetics and Molecular Ecology (5) NW
Application of molecular markers to ecology, evolution, and the management of living resources. Emphasis on understanding the strengths and weaknesses of the approach based on case studies. Prerequisite: BIOL 200. Offered: jointly with FISH 340; A.
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BIOL 350 Foundations in Physiology (3) NW
Physiology core course for biological sciences majors. Analysis of basic principles of animal and plant physiology, with emphasis on cellular processes that mediate organismic processes. Serves as gateway to upper-division courses in physiology. Prerequisite: minimum grade of 2.0 in either BIOL 220, B BIO 220, or TESC 140; recommended: either PHYS 115 or PHYS 122. Offered: AWSpS.
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BIOL 354 Foundations in Evolution and Systematics (3) NW
Evolution and systematics core course for biological sciences majors. Emphasizes patterns, processes, and consequences of evolutionary change. Serves as gateway to 400-level courses and seminars in evolution, population genetics, sociobiology, conservation biology, phylogenetics, and systematics. Prerequisite: either BIOL 180, B BIO 180, TESC 120, or BIO A 201. Offered: ASp.
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BIOL 355 Foundations in Molecular Cell Biology (3) NW
Cell biology core course for biological sciences majors. Emphasis on molecular approaches to understand cell structure, function, and regulation, and the analysis of experimental design and data interpretation. Serves as a prerequisite to advanced 400-level cell, molecular, and developmental biology courses and seminars. Prerequisite: minimum grade of 2.0 in either BIOL 200, B BIO 200, TESC 130. Offered: AWSpS.
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BIOL 356 Foundations in Ecology (3) NW
Ecology core course for biological sciences majors. Emphasizes understanding species interactions in biological communities and relationships of communities to environment. Serves as a prerequisite to 400-level courses and senior seminars in ecology, population, and conservation biology. Prerequisite: either BIOL 180, B BIO 180, or TESC 120. Offered: AW.
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BIOL 360 Cellular Anatomy (4) NW Zeman
Recognition of cellular and tissue structures in plants and animals with correlations to normal physiology and disease states. Prerequisite: minimum grade of 2.0 in either BIOL 220, B BIO 220, or TESC 140; may not be taken for credit if credit earned in BIOL 403.
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BIOL 380 Biomedical Advances and Society (3) I&S A. SCHIVELL
Recent biological advances studied in the context of our society, designed to foster critical thinking, public awareness, and policy impact. Topics may include human reproductive technologies, genetic engineering, embryonic stem cell research, and medical scanning improvements. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
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BIOL 385 Evolutionary Medicine (3) NW B. DUNCAN, D. EISENBERG
Explores evolutionary causes of health and disease. Considers how natural selection and the legacies of our human, primate, mammalian and bacterial ancestries have shaped our biology. Topics include mental disorders, aging, cancer, diet, obesity, diabetes, infectious diseases, racism, and health differences between human groups. Prerequisite: either BIO A 201 or BIOL 180. Offered: jointly with BIO A 355.
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BIOL 396 Peer Facilitation - Teaching in Biology (1-4, max. 8) NW
For undergraduates working in biology courses as peer facilitators. Peer facilitators assist with labs, lectures, and course administration and gain direct classroom experience. No independent teaching or grading. Opportunities vary by quarter and instructor. Prerequisite: either BIOL 350, BIOL 354, BIOL 355, or BIOL 356. Credit/no-credit only.
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BIOL 399 Biology Internship Program (2-12, max. 15) NW B. WIGGINS
Coordinated internship in a biology-related field. Allows a structured, real world biology work experiences off-campus. All internships must be approved by instructor. Credit/no-credit only. Offered: AWSpS.
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BIOL 400 Experiments in Molecular Biology (4) NW A. CROWE, A. SCHIVELL
Integrated reading, writing, and experimentation in molecular biology. Design and implementation of experiments using modern molecular biology techniques to address current questions in biology. Emphasizes reading and evaluating primary research literature. Includes practice in different scientific writing styles. Prerequisite: minimum grade of 2.0 in BIOL 355.
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BIOL 401 Advanced Cell Biology (3) NW
Selected topics in molecular cell biology. Strong emphasis on reading and interpreting primary research literature. Writing intensive course. Prerequisite: BIOL 355.
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BIOL 402 Functional Genomics (4) NW A. PAREDEZ
Students perform initial characterization of novel genes through epitope tagging and localization. Skills covered include: preparation of solutions, Genomic Database queries, PCR, electrophoresis, DNA purification, cloning, transformation, and immunofluorescence microscopy. Prerequisite: minimum grade of 2.0 in BIOL 355.
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BIOL 403 Physiological Mechanisms of Histology (5) NW L. ZEMAN
Develops recognition of cellular structures with correlations to normal physiology and disease states. Laboratory projects cover harvesting, sectioning, and mounting tissue. Includes weekly scientific papers. Prerequisite: either a grade of 3.0 in either BIOL 220, B BIO 220, or TESC 140, or a minimum grade of 2.0 in BIOL 350 or BIOL 355; may not be taken for credit if credit earned in BIOL 360.
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BIOL 404 Animal Physiology: Cellular Aspects (3) NW H. DE LA IGLESIA
Examines the physiology of membrane transport, nervous signaling, sensory systems, behavioral modulation, muscle, neuronal and endocrine integration, and circadian rhythms. Emphasis on the cellular and tissue level. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; either CHEM 224 or CHEM 239; either PHYS 115 or PHYS 122. Offered: A.
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BIOL 405 Cellular and Molecular Biology of Human Disease (3) NW
Emphasizes the understanding of disease mechanisms through studies of genetic and cellular basis human disease and disease models. Based on analyses of primary research articles. Prerequisite: minimum grade of 2.0 in BIOL 355; either GENOME 361 or GENOME 371; either BIOC 405, BIOC 440, BIOL 401.
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BIOL 408 Mechanisms of Animal Behavior (4) NW Beecher, Brenowitz, Sisneros
Comparative exploration of physiological and perceptual mechanisms that control behaviors necessary for survival and reproduction in animals. Model systems discussed include animal communication, mate choice, escape behavior, learning and memory, orientation, biological rhythms, foraging behavior. Prerequisite: either minimum 2.0 grade in BIOL 180, or 3.5 in PSYCH 200, or 2.0 in PSYCH 300. Offered: jointly with PSYCH 408.
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BIOL 409 Sociobiology (5) NW
Biological bases of social behavior, emphasizing evolution as a paradigm. Emphasizes how to think like an evolutionary biologist, especially with regard to interest conflict. Topics are individual versus group selection, kin selection, altruism, mating systems, sexual conflict, alternate reproductive strategies, and parent/offspring conflict. Prerequisite: either minimum grade of 2.0 in BIOL 180, 2.0 in B BIO 180, 2.0 in TESC 120, 3.5 in PSYCH 200, or 2.0 in PSYCH 300. Offered: jointly with PSYCH 409.
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BIOL 411 Developmental Biology (4) NW
Embryology and subsequent development of vertebrate and invertebrate animals, including Xenopus, mammals, chicks, Drosophila, echinoderms. Morphological changes in developing animals; experimental analysis of developing systems; underlying genetic and biochemical regulation of development. Prerequisite: minimum grade of 3.0 in either BIOL 200, B BIO 200, or TESC 130, or a minimum grade of 2.0 in BIOL 354 or BIOL 355. Offered: AW.
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BIOL 413 Molecular Genetics of Development (4) NW J. PARRISH
Uses molecular biology, cell biology, and genetic approaches to investigate how nutrient signals influence neuron growth. Prerequisite: BIOL 355.
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BIOL 414 Molecular Evolution (5) NW
Survey of empirical approaches to the study of molecular evolution and ecology, drawing on examples from a variety of taxa and the recent literature. Topics include DNA sequencing and systematics, fingerprinting approaches in behavioral ecology, and adaptive evolution at the molecular level. Prerequisite: BIOL 354. Offered: jointly with GENOME 414.
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BIOL 415 Evolution and Development (3) NW D. PARICHY
Analysis of intertwined developmental and evolutionary processes studied through evolution of developmental genes, proteins, and expression patterns in different organisms. Includes reading and analyzing implications for ecology evolution, and human disease. Prerequisite: either a minimum grade of 3.0 in BIOL 200, B BIO 200, or TESC 130, or a minimum grade of 2.0 in either BIOL 354 or BIOL 355.
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BIOL 416 Molecular Genetics of Plant Development (3)
Plant growth and development examined in molecular-genetic terms. Covers mutation, dominance, redundancy, epistasis, and key technologies for discovery of gene function as well as embryogenesis, meristem formation, flower development, and other problems in plant development. Prerequisite: minimum grade of 2.0 in BIOL 220, B BIO 220, TESC 140, GENOME 361, or GENOME 371.
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BIOL 417 Comparative Reproductive Physiology of Vertebrates (4) NW
Compares the reproductive physiology of fishes, amphibians, reptiles, birds, and mammals. Prerequisite: either BIOL 350 or BIOL 355.
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BIOL 418 Biological Clocks and Rhythms (3) NW H. DE LA IGLESIA
Examines circadian rhythms and other forms of biological rhythmicity, including annual and tidal rhythms. Includes theoretical background as well as aspects that range from the molecular and cellular basis to the ecological and evolutionary implications of biological rhythms. Prerequisite: BIOL 350 or BIOL 355.
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BIOL 421 Ecological and Evolutionary Physiology of Animals (3) NW
Explores the role of physiology in the ecology and evolution of animals. Special emphasis on how physiology influences responses to environmental change. Prerequisite: BIOL 350; either BIOL 354 or BIOL 356.
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BIOL 422 Physiology of Plant Behavior (3) NW E. VAN VOLKENBURGH
Advanced physiology seminar focusing on plant sensory mechanisms, transport and integration of information, and behavior in response to a variety of environmental stimuli. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
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BIOL 423 Marine Ecological Processes (3) NW E. CARRINGTON
Studies the ecology of the oceans and coastal regions, emphasizing benthic communities common to the Pacific Northwest. Prerequisite: either BIOL 180, B BIO 180, or TESC 120; recommended: either BIOL 250 or BIOL 356.
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BIOL 424 Plant Ecophysiology (5) NW S. KIM
Explores physiological mechanisms that underlie ecological observations, including how above- and below-ground microclimates develop and affect plant physiological processes. Discusses acclimation to environmental change along with species differences in physiological processes and plant's occupation of heterogeneous environments. Laboratories emphasize field measurement techniques. Prerequisite: either BIOL 180, B BIO 180, TESC 120, ESRM 201, ESRM 162, or FISH 162. Offered: jointly with ESRM 478; W.
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BIOL 425 Plant Physiology and Development (5) NW
Expanded coverage of plant growth, nutrition, metabolism, and development. Prerequisite: either BIOL 220, B BIO 220, or TESC 140. Offered: W.
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BIOL 427 Biomechanics (5) NW T. DANIEL
Physical biology emphasizing a mechanical approach to ecological, evolutionary, and physiological questions. Basic principles underlying fluid and solid mechanics to explore responses of animals to flows, loads, and motions. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; either MATH 125 or Q SCI 292; either PHYS 114 or PHYS 121.
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BIOL 428 Sensory Neurophysiology and Ecology (3) NW J. RIFFELL
Examines behavioral and physiological processes within an environmental framework. Uses a synthetic approach emphasizing applications to cell biology, physiology and behavior, and biomechanics. Prerequisite: either BIOL 350 or BIOL 355; either PHYS 114 or PHYS 121; recommended: linear algebra.
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BIOL 430 Marine Zoology (5) NW
Survey of groups of invertebrate animals represented in the San Juan Archipelago; natural history, functional morphology, ecology, distribution, habitat, adaptation, trophic interrelationships, and evolution. Prerequisite:BIOL 445, which must be taken concurrently; permission of Friday Harbor director. Recommended: 20 credits in biological sciences. Offered: jointly with FHL 430.
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BIOL 432 Marine Invertebrate Zoology (9) NW
Comparative morphology and biology of marine invertebrates with emphasis on field and laboratory studies. Representatives of all major and most minor phyla are collected, observed alive, and studied in detail. Taken at Friday Harbor Laboratories. Not open for credit to students who have taken BIOL 433 or BIOL 434. Recommended: 20 credits in biological sciences. Offered: S.
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BIOL 433 Marine Ecology (5) NW J. RUESINK
Study of marine ecological processes such as recruitment, disturbance, competition, and predation, and their effects on the structure and diversity of marine communities. Weekend field trips to local intertidal habitats required. Prerequisite: either BIOL 356, BIOL 472, or a minimum grade of 3.4 in either BIOL 180, B BIO 180, or TESC 120. Offered: Sp, odd years.
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BIOL 434 Invertebrate Zoology (5) NW
Comparative biology and morphology of invertebrates. Laboratory work emphasizes structures and functions. Emphasizes annelids and related worms, mollusks, and arthropods. Not open to students who have taken BIOL 430 or BIOL 432. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
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BIOL 438 Quantitative Approaches to Paleobiology, Morphology, and Systematics (5) NW
Introduction to the principles and analytical methods in the study of paleobiology, morphology, and systematics. Topics include paleobiogeography, morphology-based phylogenetics, evolutionary rates, biodiversity curves, functional morphology, morphometrics, and paleoecology. Emphasis on application of methods using fossil and modern specimens. Prerequisite: either BIOL 280, BIOL 354, or ESS 213.
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BIOL 440 General Mycology (5) NW J. AMMIRATI
General survey of the fungi with emphasis on life cycles, structure, physiology, economic importance. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
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BIOL 441 Trends in Land Plant Evolution (5) NW
Covers key innovations and trends during the evolution of land plants. Uses an evolution of development (evo-devo) approach that compares morphological, anatomical, developmental, and molecular traits across main evolutionary lineages of plants. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; either BIOL 350, BIOL 354, BIOL 355, or BIOL 356. Offered: A.
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BIOL 442 Mushrooms and Related Fungi (5) NW J. AMMIRATI
General biology, ecology, and classification of mushrooms, polypores, puffballs, and other related basidiomycetes. Emphasis on Pacific Northwest species. Prerequisite: either BIOL 360, BIOL 403, or BIOL 440.
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BIOL 443 Evolution of Mammals and their Ancestors (5) NW
Highlights the evolutionary history and systematics of mammals and their ancestors. Examines fossil and modern mammal specimens from the Burke Museum collections. Required field trip. Prerequisite: either BIOL 354, BIOL 453, or ESS 100. Offered: Sp.
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BIOL 444 Ornithology (5) NW
Field, lecture, and laboratory study of birds from an evolutionary perspective. Emphasizes taxonomy, breeding systems, brood parasitism, appearance, molt, migration, orientation, social behavior, song, and flight. Includes Saturday and weekend field trips for which students are required to share a portion of transportation costs. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; recommended: BIOL 350 or BIOL 356.
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BIOL 445 Marine Botany (5) NW
Survey of plants represented in marine environments; natural history; ecology, distribution, habitat, adaptation, and trophic interrelationships. Taken at Friday Harbor laboratories. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; BIOL 430, which must be taken concurrently. Offered: jointly with FHL 440.
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BIOL 446 Cell and Molecular Techniques in Algal Biology (3) NW R. CATTOLICO
Introduction to major algal groups emphasizing differences in form and function. Laboratory techniques include methods for culturing and plating algae; fluorescence microscopy of chloroplasts and lipid bodies; subcellular fractionation of organelles; pigment analysis and protein determination. Focus placed on scientific writing. Prerequisite: BIOL 220; BIOL 355. Offered: Sp.
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BIOL 447 The Greening of the Earth: Influence of Plants on the Evolution of Terrestrial Ecosystems (5) NW C. STROMBERG
Examines the evolution of terrestrial ecosystems focusing on how abiotic factors such as climate change have shaped the evolution of vegetation on earth, and how the evolution of vegetation has influenced the evolution of animals. Introduces paleoecological techniques using Burke Museum plant fossils. Prerequisite: either BIOL 354 or ESS 213. Offered: A.
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BIOL 448 Mammalogy (5) NW S. SANTANA MATA
Studies evolutionary patterns, ecology, behavior, physiology, and taxonomy of mammals. Relies on a comparative, systems approach to understand the major trends and mechanisms maintaining the diversity in form and function in mammals. Prerequisite: either BIOL 350, BIOL 354, or BIOL 356.
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BIOL 449 Applied Phylogenetics (3) NW A. LEACHE
Emphasizes the estimation of species trees - multilocus estimates of species or population relationships as opposed to genealogies of alleles. Provides students with the computation and bioinformatics skills needed to apply new phylogenetic techniques that can accommodate larger, more complex data sets. Prerequisite: minimum grade of 2.0 in BIOL 354.
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BIOL 450 Vertebrate Paleontology (5) NW C. SIDOR
Examines fossil vertebrate life, focusing on systematics and morphology of major lineages (fish, reptiles, bird, and early mammal relatives). Examines fossil and modern vertebrates from the Burke Museum collection in the lab. Weekend field trip. Prerequisite: either BIOL 354, BIOL 452, BIOL 453 or ESS 100. Offered: jointly with ESS 452.
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BIOL 451 Invertebrate Paleontology (5) NW
Important larger invertebrate groups; morphology, classification, stratigraphic distribution, evolution, paleoecology. Offered: jointly with ESS 451.
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BIOL 452 Vertebrate Biology (5) NW
The biology of vertebrate animals, emphasizing their diversity, adaptations, and evolutionary history. Introduces aspects of behavior, physiology, morphology, and ecology that emerge from the comparative study of vertebrates. Laboratory includes local field trips, films, and introduction to regional vertebrate fauna. Prerequisite: either BIOL 180, B BIO 180, or TESC 120. Offered: SpS.
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BIOL 453 Comparative Anatomy of Vertebrates (5) NW
Comparison of the structure of vertebrate organ systems: integument, skeletal, muscle, digestive, respiratory, cardiovascular, urinary, and reproductive, with an emphasis on evolutionary trends. Prerequisite: either BIOL 350 or BIOL 452. Offered: W.
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BIOL 454 Entomology (5) NW
Covers the biology of terrestrial arthropods, with emphasis on insects. Includes structure, classification, physiology, and ecology of insects. Interrelationships of insects and man. Requires at least one weekend field trip. Prerequisite: either BIOL 180, B BIO 180, or TESC 120.
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BIOL 457 Chemical Communication (3) NW J. RIFFELL
Exploration of how chemical signals are produced, transported, and influence behavior of microbes, plants, and animals. Synthetic approach, with emphasis on applications to cell biology, neurobiology, and ecology. Prerequisite: either a minimum grade of 2.5 in either BIOL 220, B BIO 220, or TESC 140, or a minimum grade of 2.0 in BIOL 350.
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BIOL 458 Behavioral Genetics (4) NW
Role of genetics in determining variation in human and animal behavior and in regulating behavioral development. Techniques for quantifying genetic variation, behavioral effects, and gene expression. Prerequisite: either minimum 3.5 grade in PSYCH 200, 2.0 in PSYCH 300, or 2.0 in BIOL 180. Offered: jointly with PSYCH 458.
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BIOL 459 Developmental Neurobiology (3) NW M. BOSMA
Invertebrate and vertebrate examples illustrate the mechanisms used in constructing nervous systems. Focus on the cellular and molecular mechanisms that underlie questions about the basis of neuronal diversity, axonal pathfinding and target recognition, synaptogenesis, and activity-dependent plasticity. Prerequisite: minimum grade of 2.0 in either BIOL 350 or BIOL 355. Offered: Sp.
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BIOL 460 Mammalian Physiology (3) NW
Principles of mammalian physiology with special emphasis on the cardiac, respiratory, renal systems taught at the organ and organ systems level. Prerequisite: a minimum grade of 2.5 either BIOL 350 or NBIO 301.
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BIOL 461 Neurobiology (3) NW
Broad examination of integrative mechanisms in central nervous system function, with emphasis on sensory processing, plasticity, and control of behavior. Examples are taken from a variety of animal groups. Prerequisite: minimum grade of 2.0 in BIOL 350; either PHYS 115 or PHYS 122. Offered: W.
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BIOL 462 Advanced Animal Physiology (3) NW
Physiology at levels of organisms and behavior, organ systems, and cells - an evolutionary and integrative perspective. Organismal physiology: metabolism, temperature, locomotion, osmoregulation, respiration, circulation, digestion. Prerequisite: minimum grade of 2.0 in either BIOL 350 or NBIO 301. Offered: A.
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BIOL 463 Advanced Animal Physiology Lab (3) NW
Experimental design and techniques, data analysis, written reports. Original project labs and experiments in physiology. Prerequisite: BIOL 462, which may be taken concurrently. Offered: A.
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BIOL 465 Comparative Endocrinology (3) NW
Hormonal integration of living processes at all levels in animals: molecules, cells, organs, organisms, populations. Prerequisite: BIOL 350.
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BIOL 466 Pathobiology of Emerging Diseases (3) NW L. ZEMAN
Examination of the causes, alterations in cellular function, and remediation of emerging diseases in plants and animals from a global perspective. Includes weekly scientific papers. Prerequisite: either BIOL 350, BIOL 355, BIOL 356, or BIOL 380.
View course details in MyPlan: BIOL 466

BIOL 467 Comparative Animal Physiology (3) NW
Studies organismal function in an evolutionary context. Uses a variety of animals to highlight transitions in metabolism, muscle function, respiration, circulation, digestion, excretion, and ion regulation. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
View course details in MyPlan: BIOL 467

BIOL 468 Medical Physiology (3) D. WEIGLE
Students deepen their understanding of human physiology and pathophysiology by performing in-depth analyses of actual medical cases. Cases are chosen to illustrate the mechanisms of frequently encountered diseases and the clinical presentation of affected patients. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; BIOL 350.
View course details in MyPlan: BIOL 468

BIOL 469 Evolution and Medicine (3) NW C. BERGSTROM
Responds to new applications of evolutionary biology in medicine, now being discovered at an accelerating rate. Emphasizes medically relevant aspects of evolutionary biology. Prerequisite: minimum grade of 2.0 in either BIOL 180, B BIO 180, or TESC 120.
View course details in MyPlan: BIOL 469

BIOL 470 Biogeography (4) NW
Analysis of historical and ecological determinants of current and past distributions of organisms. Integrates techniques developed by taxonomists, paleontologists, geologists, evolutionists, ecologists, and biogeographers to elucidate relationships between geographical distributions and continental drift, ecological interactions, climate, and dispersal abilities of organisms. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
View course details in MyPlan: BIOL 470

BIOL 471 Plant Ecology (5) NW
Basic concepts of plant ecology, including studies of the environment, plant-environment interactions, populations, communities, and ecosystems. Laboratory includes one weekend field trip, laboratory and greenhouse experiments, and an introduction to ecological problem solving. Prerequisite: either BIOL 180, B BIO 180, or TESC 120.
View course details in MyPlan: BIOL 471

BIOL 472 Community Ecology (5) NW
Covers the complexity of biological communities as influenced by biotic and abiotic factors, as well as the impact of human activities (like global warming) on communities. Prerequisite: BIOL 356.
View course details in MyPlan: BIOL 472

BIOL 473 Limnology (3) NW
Ecology, conservation, and management of inland aquatic ecosystems. Explores interactions among biological, chemical, and physical features of lakes and other aquatic habitats. Prerequisite: BIOL 180. Offered: jointly with FISH 473; A.
View course details in MyPlan: BIOL 473

BIOL 474 Limnology Laboratory (2) NW
Examination of biota of fresh waters, survey of limnological methods, analysis of data, and writing of scientific papers. Prerequisite: BIOL 473/FISH 473/CEE 462, which may be taken concurrently. Offered: jointly with CEE 463/FISH 474; A.
View course details in MyPlan: BIOL 474

BIOL 475 Intensive Field Experience in Biology (5) NW
Introduces field methods and research in various areas of biology, e.g., paleontology, ecology, climate change, and mycology. Includes two or more weeks away from campus at field site. Offered: S.
View course details in MyPlan: BIOL 475

BIOL 476 Conservation Biology (5) NW
Explores biological, managerial, economic, and ethical concepts affecting survival of species. Applications of ecology, biogeography, population genetics, and social sciences for the preservation of species in the face of widespread global habitat modification, destruction, and other human activities. Prerequisite: either a minimum grade of 2.5 in BIOL 180, B BIO 180, TESC 120, or BIOL 356.
View course details in MyPlan: BIOL 476

BIOL 477 Seminar in Marine Biology (3) NW Roccap
Reviews current research in marine biology. Emphasizes critical readings and discussion of primary literature. Prerequisite: FISH 250, OCEAN 250, or BIOL 250; Q SCI 381, STAT 220, or STAT 311. Offered: jointly with FISH 477/OCEAN 477; W.
View course details in MyPlan: BIOL 477

BIOL 478 Topics in Sustainable Fisheries (3, max. 9) I&S/NW
Seminar series featuring local, national, and internationally known speakers in fisheries management and conservation. Conservation/restoration in practice. Pre-seminar discussion section focusing on select readings. Topics may include harvest management, whaling, by-catch, salmon, marine protected areas, introduced species, citizen action, co-management, and marine ethics. Offered: jointly with ENVIR 478/FISH 478.
View course details in MyPlan: BIOL 478

BIOL 479 Research in Marine Biology (1-15, max. 15)
Individual research on topics in marine biology. Research projects supervised by an individual faculty member. Projects may include laboratory work, fieldwork, and literature surveys. Prerequisite: BIOL 250/FISH 250/OCEAN 250; Q SCI 381. Offered: jointly with FISH 479/OCEAN 479; AWSpS.
View course details in MyPlan: BIOL 479

BIOL 480 Field Ecology (4) NW P. BOERSMA
Field projects examining ecological and behavioral topics such as foraging and social behavior, species interactions, and structure of terrestrial and aquatic communities. Two weekend fieldtrips required. Prerequisite: either BIOL 356 or a minimum grade of 3.0 in either BIOL 180, B BIO 180, or TESC 120. Offered: Sp.
View course details in MyPlan: BIOL 480

BIOL 481 Experimental Evolutionary Ecology (5) NW B. KERR
Explores experimentally approachable questions in ecology and evolution through lectures, laboratory, and field experiments. Topics may include evolution of bacterial antibiotic resistance, the evolution of virulence, seed predation, plant biodiversity, and others. Prerequisite: either BIOL 180, B BIO 180, or TESC 120. Offered: A.
View course details in MyPlan: BIOL 481

BIOL 482 Advanced Experimental Evolutionary Ecology (2-5, max. 15) NW B. KERR
Working in pairs or independently, students pursue supervised original field or laboratory research projects. Projects span three academic quarters, with project development beginning in autumn, research continuing in winter, and culminating in spring with production of a scientific paper. Prerequisite: BIOL 481, which may be taken concurrently. Offered: WSpS.
View course details in MyPlan: BIOL 482

BIOL 483 Senior Seminar in Paleobiology (1, max. 6)
Supervised readings and group discussion. Prerequisite: either BIOL 443, BIOL 447, BIOL 450/ESS 452, BIOL 451/ESS 452, ESS 450, BIO A 388, or ARCHY 470.
View course details in MyPlan: BIOL 483

BIOL 484 Senior Seminar in Evolution and Systematics (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 354.
View course details in MyPlan: BIOL 484

BIOL 485 Senior Seminar in Cellular, Molecular and Developmental Biology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: minimum grade of 2.0 in either BIOL 350, BIOL 354, BIOL 355, BIOL 356, BIOC 405, BIOC 406, BIOC 440, BIOC 441, or BIOC 442.
View course details in MyPlan: BIOL 485

BIOL 486 Senior Seminar in Ecology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 356.
View course details in MyPlan: BIOL 486

BIOL 487 Senior Seminar in Conservation Biology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 356.
View course details in MyPlan: BIOL 487

BIOL 488 Senior Seminar in Physiology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 350.
View course details in MyPlan: BIOL 488

BIOL 489 Senior Seminar in Plant Biology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 220; recommended: one upper-division course emphasizing plant biology.
View course details in MyPlan: BIOL 489

BIOL 490 Undergraduate Seminar (1-3, max. 6) NW
Supervised readings and group discussion of selected topics of broad biological significance. Prerequisite: either BIOL 220, B BIO 220, or TESC 140.
View course details in MyPlan: BIOL 490

BIOL 492 The Teaching of Biology (2)
Basic course in the teaching of biology in the secondary school. Designed to help pre-service teachers identify useful laboratory techniques, materials, and content for the teaching of pre-college biology. Special attention to current issues in biology education. Required for biology students in Teacher Certification Program.
View course details in MyPlan: BIOL 492

BIOL 493 Study Abroad - Advanced Biology (1-15, max. 15) NW
For participants in UW Study Abroad program. Specific content varies and must be individually evaluated. Credit does not apply to major requirements without approval.
View course details in MyPlan: BIOL 493

BIOL 495 Undergraduate Research in Paleobiology (1-5, max. 10) NW
Individually arranged undergraduate research for students in the field of paleobiology.
View course details in MyPlan: BIOL 495

BIOL 497 Special Topics in Biology (1-5, max. 10) NW
View course details in MyPlan: BIOL 497

BIOL 498 Library Research (1-5, max. 10)
View course details in MyPlan: BIOL 498

BIOL 499 Undergraduate Research (1-20, max. 20)
View course details in MyPlan: BIOL 499

BIOL 500 Topics in Biology (1-3, max. 16)
View course details in MyPlan: BIOL 500

BIOL 501 Topics in Biological Instruction (1-2, max. 10)
Focused discussion of on-going research in instructional methods for life science courses. Credit/no-credit only.
View course details in MyPlan: BIOL 501

BIOL 502 Grant Writing (3)
Introduces the steps of compiling a successful grant proposal, using the NSF graduate fellowship as a template. Offered: A.
View course details in MyPlan: BIOL 502

BIOL 505 Problems in Biological Instruction (1-3, max. 10)
View course details in MyPlan: BIOL 505

BIOL 506 Scientific Manuscript Writing (4)
Introduces the writing of scientific articles. Students write a scientific manuscript, cover letter, and identify a journal for submission. Offered: W.
View course details in MyPlan: BIOL 506

BIOL 511 Topics in Mathematical Biology (1-3, max. 15)
Discussion of current topics in mathematical biology.
View course details in MyPlan: BIOL 511

BIOL 520 Departmental Seminar (1, max. 18)
Credit/no-credit only. Offered: AWSp.
View course details in MyPlan: BIOL 520

BIOL 533 Advanced Organismal Biology (9)
Advanced study and research in organismal biology. Emphasizes marine organisms and habitats of the San Juan Archipelago. Includes individual research projects. Prerequisite: permission of Director of Friday Harbor Laboratories. Offered: S.
View course details in MyPlan: BIOL 533

BIOL 536 Comparative Invertebrate Embryology (9)
Diversity in developmental patterns in major marine taxa. Analysis of evolutionary changes in development. Emphasis on observation of live embryos and larvae. Prerequisite: permission of Director of Friday Harbor Laboratories; recommended: courses in invertebrate zoology and developmental biology. Offered: SpS.
View course details in MyPlan: BIOL 536

BIOL 539 Marine Phycology (9)
Field and laboratory studies of marine algae of the San Juan Archipelago with emphasis on seaweed diversity, identification, and the role of algae in marine ecosystems. Offered at Friday Harbor Laboratories.
View course details in MyPlan: BIOL 539

BIOL 540 Seminar in Molecular, Cellular, and Developmental Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in cell, molecular, and/or developmental biology, review of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics.
View course details in MyPlan: BIOL 540

BIOL 541 Topics in Molecular, Cellular, and Developmental Biology (1-3, max. 15)
Focused discussion of on-going cell, molecular, or developmental biology research occurring in the instructor's laboratory.
View course details in MyPlan: BIOL 541

BIOL 542 Analysis of Development (1-3, max. 15)
Analysis of structural, physiological, and molecular levels of developmental processes, including gametogenesis, fertilization, cell and tissue movements, induction, and cytodifferentiation. Graduate standing or permission of instructor.
View course details in MyPlan: BIOL 542

BIOL 543 Biology of Drosophila Seminar (1, max. 12)
Weekly presentation by participants of classical literature, current literature, and research in the molecular biology, developmental biology, neurobiology, and genetics of Drosophila. Prerequisite: permission of instructor. Offered: AWSp.
View course details in MyPlan: BIOL 543

BIOL 544 Evolution of Photosynthesis (3) R. CATTOLICO
Focuses on the evolution of photosynthetic eukaryotic cells. Interactive discussions target the morphological, genetic, and biochemical consequences of progenitor cell specialization within extant taxa.
View course details in MyPlan: BIOL 544

BIOL 545 MCD-Biology Professional Skills Seminar (1, max. 6) D. PARICHY
Presentations of past or planned research in molecular, cellular, or developmental biology. Participants give presentations in either professional-meeting or chalk-talk formats. Uses written and oral evaluations by audience to focus on scientific content as well as presentation skills. Credit/no-credit only. Offered: AWSp.
View course details in MyPlan: BIOL 545

BIOL 550 Seminar in Evolution and Systematics (1-3, max. 15)
Weekly discussions of past and current scientific literature in evolution and/or systematics, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics.
View course details in MyPlan: BIOL 550

BIOL 551 Topics in Evolution and Systematics (1-3, max. 15)
Focused discussion of on-going research in evolution and/or systematics occurring in the instructor's laboratory.
View course details in MyPlan: BIOL 551

BIOL 552 Advanced Evolution ([2-5]-, max. 10)
Successful analytical approaches to understanding evolutionary patterns and the processes that generate them, examined by using a wide array of empirical and theoretical tools. Survey of how theory, modeling, and statistics can be applied to observations and experiments in evolutionary biology.
View course details in MyPlan: BIOL 552

BIOL 553 Applied Phylogenetics (3) A. LEACHE
Emphasizes the estimation of species trees - multilocus estimates of species or population relationships as opposed to genealogies of alleles. Provides students with the computation and bioinformatics skills needed to apply new phylogenetic techniques that can accommodate larger, more complex data sets. Prerequisite: BIOL 354.
View course details in MyPlan: BIOL 553

BIOL 555 Introduction to Graduate Research in Paleobiology (1)
Introduction to paleobiology techniques and resources. Credit/no-credit only. Offered: jointly with ESS 558; A.
View course details in MyPlan: BIOL 555

BIOL 557 Vertebrate Paleontology (5)
Examines the biology of vertebrate animals, emphasizing their diversity, adaptations, and evolutionary history. Introduces aspects of behavior, physiology, morphology, and ecology that emerge from the comparative study of vertebrates. Laboratory includes local field trips and introduction to regional vertebrate fauna. Offered: jointly with ESS 557.
View course details in MyPlan: BIOL 557

BIOL 560 Seminar in Ecology (1-3, max. 15)
Weekly discussions of past and current scientific literature in ecology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics.
View course details in MyPlan: BIOL 560

BIOL 561 Topics in Ecology (1-3, max. 15)
Focused discussion of on-going research in ecology occurring in the instructor's laboratory.
View course details in MyPlan: BIOL 561

BIOL 563 Experimental Evolutionary Ecology (5) B. KERR
Explores experimentally approachable questions in ecology and evolution through lectures, lab, and field experiments. Topics may include evolution of bacterial antibiotic resistance, the evolution of virulence, seed predation, plant biodiversity, and others. Corequisite: BIOL 481. Offered: A.
View course details in MyPlan: BIOL 563

BIOL 564 Advanced Experimental Evolutionary Ecology (2-5, max. 15) B. KERR
Working independently or paired with an undergraduate in BIOL 482, students pursue supervised original field or laboratory research projects. Projects span three academic quarters, with project development beginning in autumn, research continuing in winter, and culminating in spring with production of a scientific paper. Prerequisite: BIOL 563, which may be taken concurrently. Offered: AWSp.
View course details in MyPlan: BIOL 564

BIOL 565 Community Ecology (5) J. HILLE RIS LAMBERS
Covers the complexity of biological communities as influenced by biotic and abiotic factors, as wells as the impact of human activities, like global warming, on communities.
View course details in MyPlan: BIOL 565

BIOL 567 Topics in Advanced Ecology (3, max. 6)
Discusses literature on active research areas or controversies in different branches of ecology. Offered: jointly with FISH 567/SEFS 567; W.
View course details in MyPlan: BIOL 567

BIOL 570 Seminar in Conservation Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in conservation biology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. tes.
View course details in MyPlan: BIOL 570

BIOL 571 Topics in Conservation Biology (1-3, max. 15)
Focused discussion of on-going research in conservation biology occurring in the instructor's laboratory. Graduate status required, or permission of instructor for undergraduates.
View course details in MyPlan: BIOL 571

BIOL 580 Seminar in Physiology (1-3, max. 15)
Weekly discussions of past and current scientific literature in physiology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics.
View course details in MyPlan: BIOL 580

BIOL 581 Topics in Physiology (1-3, max. 15)
Focused discussion of on-going research in physiology occurring in the instructor's laboratory.
View course details in MyPlan: BIOL 581

BIOL 583 Physiological Mechanisms of Histology (5) L. ZEMAN
Develops recognition of cellular structures with correlations to normal physiology and disease states. Lab projects cover harvesting, sectioning, staining, and mounting tissue. Includes weekly scientific papers.
View course details in MyPlan: BIOL 583

BIOL 590 Seminar in Organismal Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in organismal biology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics.
View course details in MyPlan: BIOL 590

BIOL 591 Topics in Organismal Biology (1-3, max. 15)
Focused discussion of on-going research in organismal biology occurring in the the instructor's laboratory.
View course details in MyPlan: BIOL 591

BIOL 600 Independent Study or Research ([1-10]-)
Credit/no-credit only.
View course details in MyPlan: BIOL 600

BIOL 700 Master's Thesis (*-)
Credit/no-credit only. Offered: AWSpS.
View course details in MyPlan: BIOL 700

BIOL 800 Doctoral Dissertation (*-)
Credit/no-credit only. Offered: AWSpS.
View course details in MyPlan: BIOL 800