Glossary
UW Bothell Course Descriptions
UW Tacoma Course Descriptions
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Course Descriptions |
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Detailed course offerings (Time Schedule) are available for
To see the detailed Instructor Class Description, click on the underlined instructor name following the course description.
BIOL 100 Introductory Biology (5) NW
Develops an awareness of science by studying basic biological principles and their application to problems of humans and society in the contexts of special topics or themes, which vary quarter to quarter. For non-science majors only. Offered: AW.
Instructor Course Description:
Linda E. Martin-Morris
BIOL 103 Plants in the Human Environment (5) NW del Moral
Plant biology in human affairs: biodiversity, evolution, agricultural and conservation ecology, and climate change. Life cycles, plant growth and function, how plants interact with their environment and with other organisms, and how ecosystems recover from disturbance. Offered: W.
BIOL 104 Biology for Elementary School Teachers (5) NW Buttemer
Basic concepts of biology, with emphasis on background needed for confident use of the new science curriculum materials in the elementary school. Offered: AW.
Instructor Course Description:
Helen A. Buttemer
BIOL 106 Introductory Biology Seminar (1/3, max 6) NW
Focuses on current topics in biology. Topics vary from quarter to quarter. Designed to enhance learning skills of students who intend to take BIOL 161-162 or BIOL 180/200/220 and major in one of the biological sciences. Intended for HHMI Fellows. Recommended: high school chemistry and biology, or one quarter of college chemistry.
BIOL 108 Evolution and Human Behavior (3) NW
Introduction to evolution by natural selection, examining the light it can throw on human biology and behavior in such areas a s the nature of sex differences, sexual conflict, and conflict between parents and children. Does not fulfill major requirements. Offered: jointly with BIO A 100.
Instructor Course Description:
Sara L. Wyckoff
BIOL 110 Freshman Discovery Seminar in Biology (5) NW
Introduces incoming freshman to research basics and scholarly inquiry skills used in the study of biology.
BIOL 113 Introduction to Biosciences (1-3, max. 6) NW
Independent study/topics related to material taken in BIOL 180, BIOL 200, and BIOL 220. Credit/no credit only. Offered: AWSp
BIOL 114 Astrobiology: Life in the Universe (5) NW
Introduction to the new science of astrobiology, study of the origin and evolution of life on Earth, and the search for microbial and intelligent life elsewhere in the Universe. Designed for non-science, liberal arts majors. Offered: jointly with ASTR/ASTBIO/ESS/OCEAN 115.
BIOL 115 Evolution (2) NW
Evolutionary biology for nonmajors. Evolutionary history of the earth and various theories of evolution.
BIOL 117 Plant Identification and Classification (5) NW Olmstead
Plant classification and diversity of seed plants; field study and laboratory identification of the common plant families and the conspicuous flora of western and central Washington. One full-day weekend field trip (optional). For non-majors. Offered: SpS.
BIOL 118 Survey of Physiology (5) NW
Human physiology, for nonmajors and health sciences students. Offered: AWSpS.
Instructor Course Description:
Mary Pat Wenderoth
BIOL 119 Elementary Physiology Laboratory (1) NW
Prerequisite: BIOL 118 which may be taken concurrently. Offered: ASp.
BIOL 120 Current Controversies in Biology (2-5, max. 6 ) NW
Explores a current controversial topic in biology, stressing information needed by the general public to make informed personal, political, and ethical decisions relating to this topic.
BIOL 161 General Biology (5-) NW
Living systems viewed from the subcellular to the community level, emphasizing the diversity, functioning, and interaction of whole organisms. Topics covered include cell structure and function, energy, genetics, animal physiology and development. Emphasizes the position of humans in the biological world. For nonmajors and majors in biology-related fields who need a thorough two-quarter introduction to biology. Recommended: high school chemistry; high school biology. Offered: A.
BIOL 162 General Biology (5) NW
Living systems viewed from the subcellular to the community level, emphasizing the diversity, functioning, and interaction of whole organisms. Topics covered include plant and animal diversity, plant structure and function, general ecology and evolution. Emphasizes the position of humans in the biological world. For nonmajors and majors in biology-related fields who need a thorough two-quarter introduction to biology. Prerequisite: either BIOL 101 or BIOL 161. Offered: W.
BIOL 179 Learning to Learn in the Sciences (2)
Explores how to learn in the sciences. Develops various learning strategies, study skills, and personalized study plans to meet the new challenges of the BIOL 180 series through weekly in-class workshops and organized study sessions. Credit/no credit only. Corequisite: BIOL 180.
BIOL 180 Introductory Biology (5) NW
For students intending to take advanced courses in the biological sciences or enroll in preprofessional programs. Mendelian genetics, evolution, biodiversity of life forms, ecology, conservation biology. First course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Cannot be taken for credit if BIOL 203 has already been taken. Prerequisite: either CHEM 150, CHEM 152, CHEM 155, CHEM 220, CHEM 223, or CHEM 237. Offered: AWSpS.
BIOL 200 Introductory Biology (5) NW
For students intending to take advanced courses in the biological sciences or enroll in preprofessional programs. Metabolism and energetics, structure and function of biomolecules, cell structure and function, animal development. Second course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Cannot be taken for credit if BIOL 201 has already been taken. Prerequisite: 1.5 in BIOL 180; either CHEM 150, CHEM 152, CHEM 155, CHEM 220, CHEM 223, or CHEM 237. Offered: AWSpS.
BIOL 220 Introductory Biology (5) NW
For students intending to take advanced courses in the biological sciences or enroll in preprofessional programs. Animal physiology, plant development and physiology. Final course in a three-quarter series (BIOL 180, BIOL 200, BIOL 220). Cannot be taken for credit if BIOL 202 has already been taken. Prerequisite: 1.5 in BIOL 200; either CHEM 150, CHEM 152, CHEM 155, CHEM 220, CHEM 223, or CHEM 237. Offered: AWSpS.
BIOL 250 Marine Biology (3/5, max. 5) I&S, NW
Lecture-laboratory course in Marine Biology focusing on physical, biological, and social aspects of the marine environment. Topics include oceanography, ecology, physiology, behavior, conservation, fisheries, exploration, and activism. Evening marine biology movies and weekend field trip. Honors section research project. Offered: jointly with FISH/OCEAN 250.
Instructor Course Description:
M. Claire Horner-Devine
BIOL 293 Study Abroad -- Biology (1-10, max. 10) NW
For participants in UW study abroad program. Specific content varies and must be individually evaluated. Credit does not apply to major requirements without approval.
BIOL 302 Laboratory Techniques in Cell and Molecular Biology (4) NW Martin-Morris
Explores the use of various cell and molecular laboratory techniques, such as PCR, cloning, gel electrophoresis, and bacterial transformation through hands-on experiments. Students produce a portfolio of techniques they have learned. Writing credit. Prerequisite: BIOL 355. Offered: Sp.
Instructor Course Description:
Linda E. Martin-Morris
BIOL 311 Biology of Fishes (3/5) NW
Lecture and laboratory, of the morphological, physiological, behavioral, and ecological diversity of fishes of the world; designed to provide a basic foundation for advanced courses in all areas of finfish fisheries. 3-credit option does not include laboratory. Recommended: 10 credits biological science. Offered: jointly with FISH 311; W.
BIOL 317 Plant Classification and Identification (5) NW Olmstead
Classification and diversity of seed plants; concepts and principles of classification, lab and field study of common plant families in Washington, and skill development for identification of species. Cannot be taken for credit if BIOL 117 or BOTANY 113 taken. One weekend field trip. Offered: concurrently with BIOL 117; SpS.
BIOL 330 Natural History of Marine Invertebrates (5) NW
Field and laboratory course emphasizing the habits, habitats, adaptations, and interrelationships of marine animals. Weekend field trips required. Offered: S.
BIOL 331 Landscape Plant Recognition (3) NW
Field recognition of important groups of woody landscape plants, emphasizing diversity at the genus and family levels. Cultivated plant nomenclature. Plant descriptive characters evident in the field with eye and hand lens. Hardiness and landscape applications. Recommended: either BIOL 117 or BOTANY 113. Offered: jointly with ESRM 331; Sp.
BIOL 340 Genetics and Molecular Ecology (5) NW
Application of molecular markers to ecology, evolution, and the management of living resources. Emphasis on understanding the strengths and weaknesses of the approach based on case studies. Offered: jointly with FISH 340. Prerequisite: either BIOL 102, BIOL 162, or BIOL 200.
BIOL 350 Foundations in Physiology (3) NW
Physiology core course for biological sciences majors. Analysis of basic principles of animal and plant physiology, with emphasis on cellular processes that mediate organismic processes. Serves as gateway to upper-division courses in physiology. Prerequisite: either BIOL 202 or BIOL 220; recommended: either PHYS 115 or PHYS 122. Offered: AWSpS.
BIOL 354 Foundations in Evolution and Systematics (3) NW
Evolution and systematics core course for biological sciences majors. Emphasizes patterns, processes, and consequences of evolutionary change. Serves as gateway to 400-level courses and seminars in evolution, population genetics, sociobiology, conservation biology, phylogenetics, and systematics. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: WSp.
Instructor Course Description:
Billie J. Swalla
BIOL 355 Foundations in Molecular Cell Biology (3) NW
Cell biology core course for biological sciences majors. Emphasis on molecular approaches to understand cell structure, function, and regulation, and the analysis of experimental design and data interpretation. Serves as a prerequisite to advanced 400-level cell, molecular, and developmental biology courses and seminars. Prerequisite: either BIOL 200 or BIOL 201. Offered: AWSpS.
Instructor Course Description:
Merrill B Hille
BIOL 356 Foundations in Ecology (3) NW
Ecology core course for biological sciences majors. Emphasizes understanding species interactions in biological communities and relationships of communities to environment. Serves as a prerequisite to 400-level courses and senior seminars in ecology, population and conservation biology. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: AW.
BIOL 390 Undergraduate Internship Seminar (1) NW Crowe
Weekly workshop to prepare students for off-campus research internships. Required for students planning to take BIOL 399. Prerequisite: BIOL 220. Offered: AWSpS.
BIOL 396 Peer Facilitation – Teaching in Biology (1-2, max. 4) NW Martin-Morris
For students concurrently serving as peer facilitators in biology courses. Students received instruction in teaching as well as supplemental information on material they teach others. Credit no credit only. Prerequisite: either BIOL 100 or BIOL 180. Offered: AWSp.
Instructor Course Description:
Linda E. Martin-Morris
BIOL 399 Biology Internship Program (3-5, max. 15) NW Crowe
Coordinated internship in a biology-related field. Registration restricted to students already matched with an internship opportunity in BIOL 390 or by instructor approval. Students must complete at least a combined 10 credits in BIOL 399 over consecutive quarters. Credit/no credit only. Prerequisite: BIOL 390. Offered: AWSpS.
BIOL 401 Advanced Cell Biology (3) NW
Selected topics in molecular cell biology. Strong emphasis on reading and interpreting primary research literature. Writing intensive course. Prerequisite: either BIOL 355 or BIOC 442.
Instructor Course Description:
Merrill B Hille
BIOL 402 Cell Biology Laboratory (4) NW
Investigative projects using modern molecular methods (restriction enzyme digestion, blotting, hybridization, immunochemistry, density gradient centrifugation, electrophoresis) and other methods currently used to study plant and animal cells, nucleic acids, and proteins. Includes practice in scientific style writing. Prerequisite: BIOL 401, which may be taken concurrently. Offered: AWSp.
Instructor Course Description:
Alison J Crowe
Linda E. Martin-Morris
BIOL 404 Animal Physiology: Cellular Aspects (3) NW de la Inglesia
Examines the physiology of membrane transport, nervous signaling, sensory systems, behavioral modulation, muscle, neuronal and endocrine integration, and circadian rhythms. Emphasis on the cellular and tissue level. Prerequisite: BIOL 220; either CHEM 224 or CHEM 239; either PHYS 115 or PHYS 122. Offered: A.
BIOL 405 Cellular and Molecular Biology of Human Disease (3) NW
Concepts of cellular and molecular biology as applied to human disease. Emphasis on current experimental approaches to investigate disease mechanisms and the contributions of model systems. Selected topics in cancer biology, viral induced disease, gene therapy. Prerequisite: either BIOL 202 or BIOL 220; either BIOC 405, BIOC 440, BIOL 355, BIOL 401, GENET 371, or GENET 372.
BIOL 406 Insect Behavior (4) NW O'Donnell
Explores complexity and diversity of behavior in insects and related invertebrate animals. Overview of important lineages of insects and major behavioral traits. Examines how insect biology both constrains behavior and provides evolutionary opportunities. Prerequisite: either 2.0 in BIOL 180, 3.5 in PSYCH 200, or 2.0 in PSYCH 300. Offered: jointly with PSYCH 406.
BIOL 408 Mechanisms of Animal Behavior (4) NW Beecher, Brenowitz
Comparative exploration of physiological and perceptual mechanisms that control behaviors necessary for survival and reproduction in animals. Model systems discussed include animal communication, mate choice, escape behavior, learning and memory, orientation, biological rhythms, foraging behavior. Prerequisite: either 2.0 in BIOL 180, or 3.5 in PSYCH 200, or 2.0 in PSYCH 300. Offered: jointly with PSYCH 408.
BIOL 409 Sociobiology (5) NW
Biological bases of social behavior, emphasizing evolution as a paradigm. Emphasizes how to think like evolutionary biologist, especially with regard to interest conflict. Topics are individual versus group selection, kin selection, altruism, mating systems, sexual conflict, alternate reproductive strategies, and parent/offspring conflict. Prerequisite: either 3.5 in PSYCH 200, 2.0 in PSYCH 300, 2.0 in BIOL 162,or 2.0 in BIOL 180. Offered: jointly with PSYCH 409.
BIOL 411 Developmental Biology (4) NW
Embryology and subsequent development of vertebrate and invertebrate animals, including Xenopus, mammals, chicks, Drosophila, echinoderms. Morphological changes in developing animals; experimental analysis of developing systems; underlying genetic and biochemical regulation of development. Prerequisite: either BIOL 355, BIOL 401, or BIOC 442. Offered: AW.
BIOL 413 Methods and Problems in Development (3) NW Schubiger
Special topics in development. Integrating classical and current approaches. Developmental genetics, experimental embryology, molecular mechanisms of developmental regulation, and gene function in cell determination and cell differentiation in animal systems. Prerequisite: BIOL 411. Offered: irregularly.
BIOL 414 Molecular Evolution (5) NW
Survey of empirical approaches to the study of molecular evolution and ecology, drawing on examples from a variety of taxa and the recent literature. Topics include DNA sequencing and systematics, fingerprinting approaches in behavioral ecology, and adaptive evolution at the molecular level. Prerequisite: BIOL 354. Offered: jointly with GENOME 414; alternate years.
BIOL 415 Evolution and Development (3) NW Parichy, Swalla
Analysis of intertwined developmental and evolutionary processes studied through evolution of developmental genes, proteins, and expression patterns in different organisms. Includes reading and analyzing implications for ecology evolution, and human disease. Prerequisite: either BIOL 354, BIOL 355, or BIOC 442.
BIOL 416 Molecular Genetics of Plant Development (3) NW Torii
Plant growth and development examined in molecular-genetic terms. Covers mutation, dominance, redundancy, epistasis, and key technologies for discovery of gene function as well as embryogenesis, meristem formation, flower development, and other problems in plant development. Prerequisite: BIOL 220, GENOME 371.
BIOL 418 Biological Clocks and Rhythms (3) NW de la Iglesia
Examines circadian rhythms and other forms of biological rhythmicity, including annual and tidal rhythms. Includes theoretical background as well as aspects that range from the molecular and cellular basis to the ecological and evolutionary implications of biological rhythms. Prerequisite: BIOL 350 or BIOL 355.
BIOL 423 Marine Physiological Ecology (3) NW Carrington
Explores how physiology and local environment interact to influence distribution and abundance of marine organisms in a variety of habitats (rocky shore, salt marsh, coral reef), and how physiological adaptations develop. Emphasized benthic organisms that structure coastal communities (bivalves, corals, macroalgae, etc.). Prerequisite: BIOL 220; BIOL 356. Offered: alternate years.
Instructor Course Description:
Emily Carrington
BIOL 424 Plant Eco-Physiology (5) NW Ford
Explores physiological mechanisms that underlie ecological observations, including how above- and below-ground microclimates develop and affect plant physiological processes. Discusses acclimation to environmental change along with species differences in physiological processes and plant's occupation of heterogeneous environments. Laboratories emphasize field measurement techniques. Prerequisite: either BIOL 102, BIOL 162, or BIOL 220. Offered: jointly with ESRM 478.
Instructor Course Description:
Soo-Hyung Kim
BIOL 425 Advanced Plant Physiology and Development (3) NW
Expanded coverage of plant growth, nutrition, metabolism, and development. Cannot be taken for credit if BIOL 325 already taken. Prerequisite: either BIOL 102, BIOL 162, BIOL 203, or BIOL 220. Offered: W.
BIOL 426 Advanced Plant Physiology Laboratory (3) NW
Expanded laboratory experiments on the growth, nutrition, metabolism, and development of plants. Cannot be taken for credit if BIOL 326 already taken.Prerequisite: BIOL 325 or BIOL 425, which may be taken concurrently. Offered: Sp.
BIOL 427 Biomechanics (4) NW Daniel
Physical biology emphasizing a mechanical approach to ecological, evolutionary, and physiological questions. Basic principles underlying fluid and solid mechanics to explore responses of animals to flows, loads, and motions. Recommended: either BIOL 102, BIOL 162, BIOL 202, or BIOL 220; either MATH 125 or Q SCI 292; either PHYS 114 or PHYS 121.
BIOL 429 Models in Biology (4) NW
Explores use of models in biology in a wide range of topics, including morphogenesis, nerve signals, ecological interactions, population biology, and evolutionary theory. Emphasis on the biological insights models can provide rather than mathematical techniques. Prerequisite: either MATH 146, MATH 390, MATH 395, STAT 342, or STAT 391.
Instructor Course Description:
Carl T Bergstrom
BIOL 430 Marine Zoology (5) NW
Survey of groups of invertebrate animals represented in the San Juan Archipelago; natural history, functional morphology, ecology, distribution, habitat, adaptation, trophic interrelationships, and evolution. Permission of Director, Friday Harbor Laboratories required for registration. Recommended: 20 credits in biological sciences; corequisite: BIOL 445, Offered: at Friday Harbor Laboratories; Sp.
BIOL 432 Marine Invertebrate Zoology (9) NW
Comparative morphology and biology of marine invertebrates with emphasis on field and laboratory studies. Representatives of all major and most minor phyla are collected, observed alive, and studied in detail. Not open for credit to students who have taken 433 or 434. Recommended: 20 credits in biological sciences. Offered: at Friday Harbor Laboratories; S.
BIOL 433 Marine Ecology (5) NW Ruesink
Study of marine ecological processes such as recruitment, disturbance, competition, and predation, and their effects on the structure and diversity of marine communities. Weekend field trips to local intertidal habitats required. Prerequisite: either BIOL 356, BIOL 472, or a minimum grade of 3.4 in BIOL 180. Offered: Sp, odd years.
BIOL 434 Invertebrate Zoology (5) NW
Comparative biology and morphology of invertebrates. Laboratory work emphasizes structures and functions. Emphasizes annelids and related worms, mollusks, and arthropods. Not open to students who have taken 430 or 432. Prerequisite: either BIOL 102, BIOL 202, or BIOL 220.
Instructor Course Description:
Kathryn A. Mc Donald
BIOL 440 General Mycology (5) NW Ammirati
General survey of the fungi with emphasis on life cycles, structure, physiology, economic importance. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: W.
BIOL 441 Morphology and Anatomy of Land Plants (5) NW
Comparative morphology and anatomy of land plants. Derivation of morphological structures and basis for current classification schemes examined using living and fossil organisms. Laboratories emphasize live plants native to the Pacific Northwest. Prerequisite: either BIOL 102, BIOL 162, BIOL 203, or BIOL 220; recommended BIOL 317. Offered: A.
BIOL 442 Mushrooms and Related Fungi (5) NW Ammirati
General biology, ecology, and classification of mushrooms, polypores, puffballs, and other related basidiomycetes. Emphasis on Pacific Northwest species. Prerequisite: either BIOL 102, BIOL 162, BIOL 203, or BIOL 220; recommended BIOL 440. Offered: S.
BIOL 443 Evolution of Mammals and their Ancestors (5) NW
Highlights the evolutionary history and systematics of mammals and their ancestors. Examines fossil and modern mammal specimens from the Burke Museum collections. Required field trip. Prerequisite: either BIOL 354, BIOL 453, or ESS 100. Offered: A.
BIOL 444 Natural History of Birds (5) NW
Field, lecture, and laboratory study of birds framed in biological theory rather than taxonomy. Breeding systems, brood parasitism, appearance, molt, migration, orientation, social behavior, song, and flight are emphasized. Includes Saturday and weekend field trips for which students are required to share a portion of transportation costs. Prerequisite: BIOL 220; recommend BIOL 350 or BIOL 356. Offered: irregularly.
BIOL 445 Marine Botany (5) NW
Survey of plants represented in marine environments; natural history; ecology, distribution, habitat, adaptation, and trophic interrelationships. Prerequisite: either BIOL 102, BIOL 162, BIOL 203, or BIOL 220; BIOL 430/ZOOL 430, which may be taken concurrently. Offered: at Friday Harbor Laboratories; Sp.
BIOL 446 Biology of Algae (5) NW Cattolico, Waaland
Study of major algal groups emphasizing form, function, reproduction, distribution, biodiversity, and ecological roles in coastal, oceanic, and global ecosystems. Topics include classification, cellular and organisimal features, phylogeny, and evolution of major algal groups. Emphasizes economically useful and ecologically important algae. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: Sp.
BIOL 448 Marine Algal Ecology (3) NW Waaland
Marine environment in relation to requirements for algal life. Intertidal, subtidal, geographical, and seasonal distribution of marine algae. Functional aspects of algal form, structure, productivity, and energy economy of marine algal communities. Algal utilization and aquaculture. Offered: A.
BIOL 450 Vertebrate Paleontology (4) NW Sidor
Examines vertebrate life as seen in the fossil record, focusing first on fish lineages, and then on amniote (reptile, bird, and early mammal relatives) systematics and morphology. Examines fossil and modern vertebrates from the Burke Museum collection in the lab. One required weekend field trip. Prerequisite: either BIOL 453 or ESS 100. Offered: jointly with ESS 452; alternate years.
BIOL 451 Invertebrate Paleontology (5) NW Ward
Important larger invertebrate groups; morphology, classification, stratigraphic distribution, evolution, paleoecology. Offered: jointly with ESS 451; W.
BIOL 452 Vertebrate Biology (5) NW
The biology of vertebrate animals, emphasizing their diversity, adaptations, and evolutionary history. Introduces aspects of behavior, physiology, morphology, and ecology that emerge from the comparative study of vertebrates. Laboratory includes local field trips, films, and introduction to regional vertebrate fauna. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or both BIOL 202 and BIOL 203. Offered: SpS.
BIOL 453 Comparative Anatomy of Vertebrates (5) NW
Comparison of the structure of vertebrate organ systems: integument, skeletal, muscle, digestive, respiratory, cardiovascular, urinary, and reproductive, with an emphasis on evolutionary trends. Prerequisite: BIOL 350. Offered: W.
BIOL 454 Entomology (3) NW
Biology of terrestrial arthropods, with emphasis on insects. Structure, classification, physiology, and ecology of insects. Interrelationships of insects and man. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or both BIOL 202 and BIOL 203.
BIOL 455 Entomology Laboratory (3) NW
Structure and function of arthropods, with emphasis on insects. Field studies and taxonomy of important insect groups. Students may be required to share a portion of the transportation costs of field trips. Prerequisite: BIOL 454 which may be taken concurrently. Offered: Sp.
BIOL 456 Vegetation of Western Washington (5) NW del Moral
Vegetation of western Washington, including mature, seral, and weedy vegetation. Recognition of landscape patterns, sight identification of common and indicator species, classification of major community types, and uses of native species in landscape design. Four weekend field trips required. Prerequisite: either BIOL 117 or BIOL 317.
BIOL 458 Behavioral Genetics (4) NW O'Donnell
Role of genetics in determining variation in human and animal behavior and in regulating behavioral development. Techniques for quantifying genetic variation, behavioral effects, and gene expression. Prerequisite: either 3.5 in PSYCH 200, 2.0 in PSYCH 300, or 2.0 in BIOL 161, or 2.0 in BIOL 180. Offered: jointly with PSYCH 458.
BIOL 459 Developmental Neurobiology (3) NW Bosma
Invertebrate and vertebrate examples illustrate the mechanisms used in constructing nervous systems. Focus on the cellular and molecular mechanisms that underlie questions about the basis of neuronal diversity, axonal pathfinding and target recognition, synaptogenesis, and activity-dependent plasticity. Prerequisite: BIOL 220. Offered: Sp.
BIOL 460 Mammalian Physiology (3) NW
Principles of mammalian physiology with special emphasis on the cardiac, respiratory, renal systems taught at the organ and organ systems level. Prerequisite: BIOL 350.
Instructor Course Description:
Mary Pat Wenderoth
BIOL 461 Neurobiology (3) NW Bosma, Perkel
Broad examination of integrative mechanisms in central nervous system function, with emphasis on sensory processing, plasticity, and control of behavior. Examples are taken from a variety of animal groups. Prerequisite: BIOL 220; either PHYS 115 or PHYS 122. Offered: W.
BIOL 462 Advanced Animal Physiology (3) NW Huey, Wenderoth
Physiology at levels of organisms and behavior, organ systems, and cells -- an evolutionary and integrative perspective. Organismal physiology: metabolism, temperature, locomotion, osmoregulation, respiration, circulation, digestion. Prerequisite: either BIOL 202 or BIOL 220; either BIOL 350 or BIOL 460; either CHEM 152, CHEM 155, CHEM 160, CHEM 162, CHEM 164, CHEM 165, or CHEM 220; either PHYS 114 or PHYS 121. Offered: A.
BIOL 463 Advanced Animal Physiology Lab (3) NW Huey, Wenderoth
Experimental design and techniques, data analysis, written reports. Original project labs and experiments in physiology. Prerequisite: BIOL 462, which may be taken concurrently. Offered: A.
Instructor Course Description:
Mary Pat Wenderoth
BIOL 470 Biogeography (4) NW
Analysis of historical and ecological determinants of current and past distributions of organisms. Integrates techniques developed by taxonomists, paleontologists, geologists, evolutionists, ecologists, and biogeographers to elucidate relationships between geographical distributions and continental drift, ecological interactions, climate, and dispersal abilities of organisms. Not available for credit if credit has previously been given for ZOOL 475. Recommended: one year college biology; background in ecology and evolution. Offered: irregularly.
BIOL 471 Plant Ecology (5) NW del Moral
Basic concepts of plant ecology, including studies of the environment, plant-environment interactions, populations, communities, and ecosystems. Laboratory includes one weekend field trip, laboratory and greenhouse experiments, and an introduction to ecological problem solving. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: alternative years; Sp.
BIOL 472 Community Ecology (4) NW
Explores the complexity of biological communities and interactions of plant, animal, and microbes in marine, freshwater, and terrestrial ecosystems. Promotes synthetic and integrative thinking in the environmental sciences through the study of theoretical and empirical issues and original research data. Prerequisite: BIOL 356; recommended: solid foundation in math and physics.
BIOL 473 Limnology (3) NW Schindler
Ecology, conservation and management of inland aquatic ecosystems. Explores interactions among biological, chemical and physical features of lakes and other aquatic habitats. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203. Offered: jointly with FISH 473; A.
BIOL 474 Limnology Laboratory (2) NW Schindler
Examination of biota of fresh waters, survey of limnological methods, analysis of data, and writing of scientific papers. Prerequisite: BIOL 473/FISH 473/CEE 462, which may be taken concurrently. Offered: jointly with FISH 474/CEE 463; A.
BIOL 476 Conservation Biology (5) NW Boersma, Tewksbury
Explores biological, managerial, economic, and ethical concepts affecting survival of species. Applications of ecology, biogeography, population genetics, and social sciences for the preservation of species in the face of widespread global habitat modification, destruction, and other human activities. Prerequisite: either BIOL 102, BIOL 162, BIOL 180, or BIOL 203.
BIOL 478 Topics in Sustainable Fisheries (3, max. 9) I&S/NW Parrish
Seminar series featuring local, national and internationally known speakers in fisheries management and conservation. Case studies. Conservation/restoration in practice. Pre-seminar discussion section focusing on select readings. Final paper. Topics may include harvest management, whaling, by-catch, salmon, marine protected areas, introduced species, citizen action, co-management, and marine ethics. Offered: jointly with ENVIR/FISH 478; odd years; W.
BIOL 480 Field Ecology (4) NW Boersma
Field projects examining ecological and behavioral topics such as foraging and social behavior, species interactions, and structure of terrestrial and aquatic communities. Two weekend fieldtrips required. Prerequisite: either BIOL 356 or BIOL 472. Offered: Sp.
BIOL 481 Experimental Evolutionary Ecology (5) NW Bradshaw, Kerr, Tewksbury
Explores experimentally approachable questions in ecology and evolution through lectures, lab, and field experiments. Topics may include evolution of bacterial antibiotic resistance, the evolution of virulence, seed predation, plant biodiversity, and others. Prerequisite: BIOL 180. Offered: A.
Instructor Course Description:
Joshua R Nahum
BIOL 482 Advanced Experimental Evolutionary Ecology (2-5, max, 15) NW Bradshaw, Kerr, Tewksbury
Working in pairs or independently, students pursue supervised original field or lab research projects. Projects span three academic quarters, with project development beginning in autumn, research continuing in winter, and culminating in spring with production of a scientific paper. Prerequisite: BIOL 481, which may be taken concurrently. Offered: AWSp.
BIOL 484 Senior Seminar in Evolution and Systematics (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 354.
BIOL 485 Senior Seminar in Cellular, Molecular and Developmental Biology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 355.
Instructor Course Description:
Merrill B Hille
BIOL 486 Senior Seminar in Ecology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 356.
Instructor Course Description:
Adelaide C. E. Rhodes
BIOL 487 Senior Seminar in Conservation Biology (1-3, max. 9) NW
Supervised readings and group discussion. Recommended: one upper division course in ecology or conservation biology.
Instructor Course Description:
Martha Groom
BIOL 488 Senior Seminar in Physiology (1-3, max. 9) NW
Supervised readings and group discussion. Prerequisite: BIOL 350.
BIOL 489 Senior Seminar in Plant Biology (1-3, max. 9) NW
Supervised readings and group discussion. Recommended: one upper division course emphasizing plant biology
Instructor Course Description:
Veronica Sandra Distilio
BIOL 490 Undergraduate Seminar (1-3, max. 6) NW
Supervised readings and group discussion of selected topics of broad biological significance. Prerequisite: BIOL 102, BIOL 162, BIOL 203, or BIOL 220.
Instructor Course Description:
Caroline A Stromberg
BIOL 491 Special Topics in Biological Science for Teachers (1-9, max. 9) NW
Study of selected areas of biology. Designed to enhance the skills and background of K-12 teachers. Credit/no credit only. Recommended: teaching experience.
BIOL 492 The Teaching of Biology (2)
Basic course in the teaching of biology in the secondary school. Designed to help preservice teachers identify useful laboratory techniques, materials, and content for the teaching of pre-college biology. Special attention to current issues in biology education. Required for biology students in Teacher Certification Program.
BIOL 493 Study Abroad -- Advanced Biology (1-15, max. 15) NW
For participants in UW study abroad program. Specific content varies and must be individually evaluated. Credit does not apply to major requirements without approval.
BIOL 496 Peer Teaching Assistants in Biology (1-5, max. 10)
Direct experience in the classroom, typically teaching a lab section of BIOL 100. Peer Teaching Assistants attend lectures and weekly preparation meetings and gain in-depth background on the subject material as well as training in teaching techniques and approaches. Credit/no credit only. Prerequisite: either BIOL 102, BIOL 162, BIOL 220, or both BIOL 202 and BIOL 203. Offered: AWSp.
Instructor Course Description:
Linda E. Martin-Morris
Mary E Nicotri
BIOL 497 Special Topics in Biology (1-5, max. 10) NW
BIOL 498 Library Research (1-5, max. 10)
BIOL 499 Undergraduate Research (1-20, max. 20)
BIOL 500 Topics in Biology (1-3, max. 16)
BIOL 505 Problems in Biological Instruction (1-3, max. 3)
Instructor Course Description:
Martha Groom
BIOL 510 Seminar in Mathematical Biology (1-3, max. 15) Bergstrom, Daniel, Grunbaum, Kot, Odell, Thompson
Seminar on a topic of current interest in mathematical biology. Graduate status required, or permission of instructor for undergraduates.
BIOL 511 Topics in Mathematical Biology (1-3, max. 15)
Discussion of current topics in mathematical biology. Graduate status required, or permission of instructor for undergraduates.
BIOL 520 Departmental Seminar (1, max. 18)
Credit/no credit only. Offered: AWSp.
BIOL 533 Advanced Invertebrate Zoology (9)
Invertebrate fauna of the San Juan Archipelago. Topic changes from year to year. Individual research projects are emphasized. Prerequisite: course in invertebrate zoology and permission of Director of Friday Harbor Laboratories. Offered: at Friday Harbor Laboratories; SpS.
BIOL 536 Comparative Invertebrate Embryology (9)
Diversity in developmental patterns in major marine taxa. Analysis of evolutionary changes in development. Emphasis on observation of live embryos and larvae. Prerequisite: permission of Director of Friday Harbor Laboratories; recommended: courses in invertebrate zoology and developmental biology. Offered: at Friday Harbor Laboratories; SpS.
BIOL 539 Marine Phycology (9)
Field and laboratory studies of marine algae of the San Juan Archipelago with emphasis on seaweed diversity, identification, and the role of algae in marine ecosystems. Offered at Friday Harbor Laboratories.
BIOL 540 Seminar in Molecular, Cellular, and Developmental Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in cell, molecular, and/or developmental biology, review of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Graduate status required, or permission of instructor for undergraduates.
BIOL 541 Topics in Molecular, Cellular, and Developmental Biology (1-3, max. 15)
Focused discussion of on-going cell, molecular, or developmental biology research occurring in the instructor's laboratory. Graduate status required, or permission of instructor for undergraduates.
BIOL 542 Analysis of Development (1-3, max. 15)
Analysis of structural, physiological, and molecular levels of developmental processes, including gametogenesis, fertilization, cell and tissue movements, induction, and cytodifferentiation. Graduate standing or permission of instructor.
BIOL 543 Biology of Drosophila Seminar (1, max. 12)
Weekly presentation by participants of classical literature, current literature, and research in the molecular biology, developmental biology, neurobiology, and genetics of Drosophila. Prerequisite: permission of instructor. Offered: AWSp.
BIOL 546 Experimental Design in Cell Biology (1.5) Wakimoto, Wright, Hille, Cooper
Focuses on experimental design in cell biology. A topic of current research interest is covered in depth in order to follow a line of investigation and critically evaluate the strengths and limitations of various experimental strategies. Offered: jointly with CONJ 536; W.
BIOL 550 Seminar in Evolution and Systematics (1-3, max. 15)
Weekly discussions of past and current scientific literature in evolution and/or systematics, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Graduate status required, or permission of instructor of undergraduates.
BIOL 551 Topics in Evolution and Systematics (1-3, max. 15)
Focused discussion of on-going research in evolution and/or systematics occurring in the instructor's laboratory. Graduate status required, or permission of instructor for undergraduates.
Instructor Course Description:
Veronica Sandra Distilio
BIOL 552 Advanced Evolution (3)
Successful analytical approaches to understanding evolutionary patterns and the processes that generate them, examined by using a wide array of empirical and theoretical tools. Survey of how theory, modeling, and statistics can be applied to observations and experiments in evolutionary biology. Graduate standing required.
BIOL 560 Seminar in Ecology (1-3, max. 15)
Weekly discussions of past and current scientific literature in ecology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Graduate status required, or permission of instructor for undergraduates.
Instructor Course Description:
Joshua J Tewksbury
BIOL 561 Topics in Ecology (1-3, max. 15)
Focused discussion of on-going research in ecology occurring in the instructor's laboratory. Graduate status required, or permission of instructor for undergraduates.
BIOL 562 Advanced Ecology (3)
Successful analytical approaches to understanding ecological patterns and the mechanisms that generate them, examined by using a wide array of empirical and theoretical tools. Applying theory, modeling, and statistics to empirically derived data to providing insight and solutions to key environmental problems. Graduate standing required.
BIOL 563 Experimental Evolutionary Ecology (5) NW Bradshaw, Kerr, Tewksbury
Explores experimentally approachable questions in ecology and evolution through lectures, lab, and field experiments. Topics may include evolution of bacterial antibiotic resistance, the evolution of virulence, seed predation, plant biodiversity, and others. Offered: A; concurrent with BIOL 481.
BIOL 564 Advanced Experimental Evolutionary Ecology (2-5, max, 15) NW Bradshaw, Kerr, Tewksbury
Working independently or paired with an undergraduate in BIOL 482, students pursue supervised original field or lab research projects. Projects span three academic quarters, with project development beginning in autumn, research continuing in winter, and culminating in spring with production of a scientific paper. Prerequisite: BIOL 563, which may be taken concurrently. Offered: AWSp.
BIOL 570 Seminar in Conservation Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in conservation biology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Graduate status required, or permission of instructor for undergraduates.
Instructor Course Description:
Martha Groom
BIOL 571 Topics in Conservation Biology (1-3, max. 15)
Focused discussion of on-going research in conservation biology occurring in the instructor's laboratory. Graduate status required, or permission on the instructor for undergraduates.
Instructor Course Description:
Martha Groom
BIOL 572 Science and Environmental Policy (3-5, max. 5) Boersma
Instructor Course Description:
P Dee Boersma
BIOL 580 Seminar in Physiology (1-3, max. 15)
Weekly discussions of past and current scientific literature in physiology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Graduate status required, or permission of instructor for undergraduates.
BIOL 581 Topics in Physiology (1-3, max. 15)
Focused discussion of on-going research in physiology occurring in the instructor's laboratory. Graduate status required, or permission of instructor for undergraduates.
BIOL 590 Seminar in Organismal Biology (1-3, max. 15)
Weekly discussions of past and current scientific literature in organismal biology, reviews of the state of the field, and presentation of research results. Discussions may cover the full breadth of the discipline or focus on selected topics. Prerequisite: graduate standing, or permission of instructor for undergraduates.
Instructor Course Description:
Veronica Sandra Distilio
Joshua J Tewksbury
BIOL 591 Topics in Organismal Biology (1-3, max. 15)
Focused discussion of on-going research in organismal biology occurring the the instructor's laboratory. Prerequisite: graduate standing, or permission of instructor for undergraduates.
BIOL 600 Independent Study or Research (1-10)
Credit/no credit only.
BIOL 700 Master's Thesis (*)
Credit/no credit only. Offered: AWSpS.
BIOL 800 Doctoral Dissertation (*)
Credit/no credit only. Offered: AWSpS.
